Showing posts with label France. Show all posts
Showing posts with label France. Show all posts

Friday, 13 June 2025

Malformations in Trilobites from the Silurian and Devonian of Europe.

Malformations in fossils, such as pathologies caused by infections, scars left by recovered injuries, or teratologies caused by developmental problems, can tell us a lot about how extinct organisms grew and interacted with their environments, although when analysing these is clearly preferable to have access to non-malformed examples of the same species, or at least a close relative.

Trilobites were a diverse and abundant group of Arthropods which appeared early in the Cambrian, and survived until the End Permian Extinction. Their dorsal exoskeleton was heavily biomineralized, and was shed periodically to allow the animals to grow. This has lead to an extensive fossil record with Trilobites being extremely common in many Palaeozoic marine deposits. This abundant fossil record makes Trilobites an excellent candidate group for the study of malformations.

In a paper published in the journal Acta Palaeontologica Polonica on 22 April 2025, Russell Bicknell of the Division of Paleontology (Invertebrates) at the American Museum of Natural History, and the Palaeoscience Research Centre at the University of New EnglandPatrick Smith of the Palaeontology Department at the Australian Museum Research Institute, and the Department of Biological Sciences at Macquarie UniversityLisa Amati of Paleontology at the New York State Museum, and Melanie Hopkins, also of the Division of Paleontology (Invertebrates) at the American Museum of Natural History, describe malformations in European Silurian and Devonian Trilobite specimens from the collections of the Natural History Museum in London and the New York State Museum.

The first specimen examined by Bicknell et al., NYSM 19739, is an isolated cephalon (head part) from a Harpetid Trilobite, Lioharpes venulosus, from the Early Devonian Koněprusy Limestone of the Czech Republic, in the collection of the New York State Museum. The cephalon is 26.2 mm long and 22.3 mm wide with a u-shaped indentation on its right marginal rim. This indentation is 5.6 mm long and extends 1.9 mm towards the midline. The marginal rim is covered in small circular pits, which around the indentation are irregular, ovate, and occasionally fused into larger pits.

Malformed Harpetid Trilobite Lioharpes venulosus, NYSM 19739 from the Koněprusy Limestone, Pragian, Lower Devonian, Koněprusy, Czech Republic. (A₁) complete cephalon; (A₂) close up showing U-shaped indentation (arrow). Specimen coated in ammonium chloride sublimate. Bicknell et al. (2025).

Bicknell et al. note that malformations to the cephalic fringes of Harpetid Trilobites have been recorded before, and that these are usealy attributed to injuries, an analysis with which they concur. However, they also observe that injuries can happen in a variety of ways, with fringe injuries having previously attributed to problems during moulting, failed predation attempts, or unknown causes. They suggest that a moulting injury is the most likely explanation for the injury to the Koněprusy specimen, with the delicate fringe likely torn during moulting, and the enlarged and fused pits being a result of fusion of the torn margin during healing. Various purposes have been suggested for the cephalic fringes of Harpetid Trilobites, including filtering for food, sensory roles, sediment ploughing, hydrostatic support, cephalic reinforcement, burrowing, and enhancing hydrodynamic efficiency. Whatever the purpose of this organ, an injury to it is likely to have been detrimental to the living Trilobite, and presumably repairing this injury would have been a priority during subsequent moults.

The second specimen, NHMUK PI In 65061, looked at is a Phacopid Trilobite, Calymene blumenbachii, from the Early Silurian Much Wenlock Limestone Formation of Shropshire, England, in the collection of the Natural History Museum. This specimen comprises a partial cephalon, thorax, and pygidium, with a total length of 92.9 mm and a width of 48.6 mm. The second thoracic axial ring (middle part of the second segment of the thorax) of this specimen is covered by a structure with closely spaced openings, which has an elevated round crater at its right extremity, with an opening 1.7 mm across.

Bicknell et al. interpret this as an encrusting Trepostome Bryozoan covering the 3rd thoracic tergite, with the larger opening being an ovate zoarium (specially modified zooid which produced eggs). The restriction of the encrustation to one tergite strongly suggests that this happened while the Trilobite was alive, and that the Bryozoan colony was therefore unable to overgrow the articulations between tergites.

Abnormal Calymenid Trilobites Calymene blumenbachii from the Much Wenlock Limestone Formation, Homerian, Wenlock, Silurian, England, UK. (A) NHMUK PI In 65061, (A₁) complete specimen; (A₂) close up showing the large bryozoan growth. (B) NHMUK PI In 19857 showing pygidial ribs that terminate early (white arrows) and are fused proximal to the medial lobe (black arrow).  Bicknell et al. (2025).

The next specimen examined, NHMUK PI In 19857, is another example of Calymene blumenbachii from the Much Wenlock Limestone. This specimen is a partial pygidium (tail part) 13.2 mm long and 18.2 mm wide. On the right side of this specimen the pygidial ribs are disrupted and irregular, with two ribs terminating 1.6 mm short of the pygidial margin, while another two fuse 1.2 mm from the pygidial axis.

Bicknel et al. observe that similar deformations to the pygidia have been observed in a wide range of other Trilobites, including Dalmanities pleuroptyxDechenella macrocephalusNiobina sp., and Prionopeltis archiaci. They attribute these deformations to genetic or developmental issues, but suggest that the limited disruption to the pygidium they caused did not represent a major handicap.

Specimen NYSM 19740 is an Acastid Trilobite, Treveropyge sp., from the early Devonian Saint Céneré Formation of Mayenne in northwest France. this specimen is another isolated pygidium, s 11.6 mm long and 17.9 mm wide. It has a deformation to the axial lobe, which is asymmetric, with two of the axial rings malformed and curving to the right.

Malformed Acastid Trilobite Treveropyge sp., NYSM 19740 from the Saint Céneré Formation, Lochkovian, Lower Devonian, Mayenne, France. (A₁) Complete pygidium; (A₂) close up showing asymmetrical axial lobe and incomplete axial ring (arrows). Specimen coated in ammonium chloride sublimate. Bicknell et al. (2025).

Again, Bicknell et al. note that similar deformations have been seen in other Trilobites, such as Calliops marginatusDolicholeptus licticallis, and Sceptaspis lincolnensis. They suggest that these malformations are caused by genetic deformations, leading to incomplete development or non-functional somites.

The final specimen examined, NHMUK PI I 1108, is an external impression of a partial pygidium belonging to the Styginid Trilobite Scutellum (Scutellum) pardalios, from the Middle Devonin Barton Limestone Member of Devon, England. This impression is t is 59.5 mm long and 44.0 mm wide. On the right side of this impression (i.e. on the left side of the Trilobite) two ribs fuse into a single rib 29.1 mm from the pygidial axis. This single rib then terminates 4.8 mm from the pygidial margin.

Malformed Styginid Trilobite Scutellum (Scutellum) pardalios, NHMUK PI I 1108 from the Barton Limestone Member, Torquay Limestone Formation, Givetian, Middle Devonian, England, UK. (A₁) Pygidium preserved as external impression; (A₂) close up showing fused pygidial pleurae (arrows). Bicknell et al. (2025).

Bicknell et al. note that the surface of the pygidium was covered with ornamentation, with no visible break in this, which appears to  rule out the malformation having been formed by an unsuccessful predation attempt, or similar injury. Recovery from injury is the most commonly sited reason for malformations seen in Styginid Trilobites, followed by parasitic infections during early development, which seems equally unlikely. Bicknell et al. suggest instead that this deformity might be the result of a difficult moult, or possibly a genetic aberration. They do not believe this minor deformity would have significantly affected the living Trilobite.

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Friday, 11 April 2025

Fireball meteor over Luxembourg.

Witnesses across Belgium, the Netherlands, Luxembourg, Switzerland, northern France, western Germany, southeast England, northern Italy, and parts of Austria, have reported observing a bright fireball meteor around 2.30 am local time (around 0.30 am GMT) on Sunday 6 April 2025. The fireball is described as having moved from southeast to northwest, appearing over the Grand Est region of northeastern France and disappearing overr Luxembourg. A fireball is defined as a meteor (shooting star) brighter than the planet Venus. These are typically caused by pieces of rock burning up in the atmosphere, but can be the result of man-made space-junk burning up on re-entry.

The 6 April 2025 meteor seen from the Saint-Luperce, France. Romain Kropp/Météo-Centre-Val de Loire/American Meteor Society.

Objects of this size probably enter the Earth's atmosphere several times a year, though unless they do so over populated areas they are unlikely to be noticed. They are officially described as fireballs if they produce a light brighter than the planet Venus. The brightness of a meteor is caused by friction with the Earth's atmosphere, which is typically far greater than that caused by simple falling, due to the initial trajectory of the object. Such objects typically eventually explode in an airburst called by the friction, causing them to vanish as a luminous object. However, this is not the end of the story as such explosions result in the production of a number of smaller objects, which fall to the ground under the influence of gravity (which does not cause the luminescence associated with friction-induced heating).

Heat map showing areas where sightings of the meteor were reported (warmer colours indicate more sightings), and the apparent path of the object (blue arrow). American Meteor Society.

These 'dark objects' do not continue along the path of the original bolide, but neither do they fall directly to the ground, but rather follow a course determined by the atmospheric currents (winds) through which the objects pass. Scientists are able to calculate potential trajectories for hypothetical dark objects derived from meteors using data from weather monitoring services.

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Tuesday, 21 January 2025

Series of 'curse-amulets' found in burials at Romano-Gaulish cemetery in Orléans, France.

Archaeologists from Le Service Archaeologie Orléans have discovered a series of  'curse-amulets' in burials at a Romano-Gaulish cemetery beneath an eighteenth century hospital in the city of Orléans in north-central France, according to a press release issued on 7 January 2025. The cemetery, which was discovered in 2022, was already considered unusual, in that it comprises a single row of about 60 graves along a wall, and that all of those buried here appear to have been adult males, contrary to typical Roman practice. The burials are thought to date from the first-to-third centuries AD. All of the dead were buried in painted wooden coffins, with no sign of any cremations. Curiously, 21 of the graves also contained 'curse-amulets'.

Location of the 'curse-amulet', labelled as 'Tablette de défixion' within burial F2199 at the Orléans Romano-Gaulish cemetery. Le Service Archaeologie Orléans.

'Curse-amulets' (properly called 'defixion tablets') are sheets of lead upon which an inscription calling for supernatural intervention has been written. The use of lead was not just practical (although it was practical, since the soft nature of lead made it easy to inscribe and fold or roll up) but also served a symbolic purpose, lead being associated with the underworld and the gods who dwelt there in the Greek and Roman worlds. Thus, the invocations on these amulets were typically calling on these gods to cause harm to someone, with more benign prayers written on different materials. Curse amulets were often hidden somewhere on their intended victims property, or buried deep in the ground where they might better reach the chthonic deities.

The curse amulet recovered from grave F2199 in its folded state. Le Service Archaeologie Orléans.

To date, one of the curse amulets from Orléans has been partially deciphered. This amulet, from grave F2199, was carefully unrolled and then chemically treated to remove patches of corrosion and protect its surface from further degradation. The inscription thus revealed was to faint to be read by the Human eye, but has been largely revealed using Reflectance Transformation Imaging, a method which involves taking photographs from a large number of different angles, then using software to create a three-dimensional model of the surface. An online tool showing Reflectance Transformation Imaging of the amulet can be found here

The text revealed on the amulet from grave F2199 by Reflectance Transformation Imaging. Le Service Archaeologie Orléans.

The inscription revealed appears to be in the ancient Gallic language, written in Latin cursive, with some borrowed words from Greek and Latin. This has been interpreted as 'ib r…mi [m]arte rigisamu | a]nmantigIu, Se uiron bnanon uanderonado brixton sod-esti, Cisin…piSSlon atlon atemiston, Etic se-uiron banon canti piSSiantas, Sollebne(m), Marulliam, Sulpici(i), Claudia(m), Marulliam g., Curiatiu(m), Mat|(e)rno(n), Tiberium, Cantognati, Sulpici(i), (B)regesia, Regina Italica pri(uata) Sulpici(i), Regina Regina, Regina dona Tibe(rius), [..]ix, Ateporigis, Lecti, (Se)gouisu(m)'. 

Gallic is not a well-understood language, but can be deciphered to some extent by comparison to other ancient Celtic languages, such as Old Irish, Old Breton, and Old Welsh. The inscription  has been translated by the linguist Pierre-Yves Lambert as reading 'To Mars the Royal, who pierces names, It is the bewitchment of these men and women below (named) who accomplished the unfortunate and unjust feat, and also all those who were accomplices of these men and women', followed by a list of Latin names, then an invocation to 'Regina' (presumably a reference to a deity), then some more names in Gallic.

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Sunday, 24 November 2024

Numenius tenuirostris: The Slender-billed Curlew declared extinct.

The Earth's biodiversity is considered to be facing a crisis at a global level, with many experts believing that species may be going extinct at a rate as high as during the great extinctions recorded in the fossil record. Despite this, it is very hard to determine how many species are going extinct, in part because many species have never been documented, but also because it is generally impossible to tell whether a species is extinct, unless it can be confidently asserted that all populations were being monitored prior to extinction. The best recorded organisms tend to be Vertebrates, and one of the most extensively monitored Vertebrate groups are the Birds, a group which tend to be highly visible, and which are often recorded extensively by non-professional citizen-science groups. Despite this attention, many species of Birds have not been seen for a long time, and this cannot always be taken as evidence that they have become extinct: for instance, the Black-browed Babbler, Malacocincla perspicillata, was rediscovered in 2020 after not being observed in 180 years. The apparent extinction of species can be problematic, causing conservationists to cease efforts to protect a species which is close to extinction; conversely failure to realise that a species has become extinct can lead to efforts being dedicated to preserving it which could otherwise have been directed towards other species which might still be saved.

In a paper published in the journal Ibis on 17 November 2024, Greame  Buchanan of the RSPB Centre for Conservation Science, Ben Chapple of the Centre for Biodiversity and Environment Research at University College London, Alex Berryman of BirdLife International, Nicola Crockford of the Royal Society for the Protection of Birds, Justin Jansen of the Naturalis Biodiversity Center, and Alexander Bond of the Bird Group at the Natural History Museum, formally declare the Slender-billed Curlew, Numenius tenuirostris, to be extinct.

The International Union for the Conservation of Nature's Red List of Threatened Species currently lists the Slender-billed Curlew as Critically Endangered, on the assumption that the total population is less than 50 Birds and declining. The species is believed to be restricted to the Palaearctic biogeographical region, breeding in central Asia and migrating to Europe, North Africa, the Middle East, and the Arabian Peninsula. Slender-billed Curlews are thought to breed to the east of the Ural Mountains, in the area around Omsk in southern Russia. Isotope studies of museum specimens suggest that the species may also have bred further south, in northern Kazakhstan, while some eggs assigned to the species, also in museum collections, indicate that the species may also have bred to the west of the Urals. Outside of the breeding season the Birds ranged west as far as Western Europe and the Atlantic coast of North Africa, being known from across the Mediterranean Region, the Middle East, the Arabian Peninsula, and the Pannonian Plains of southeast Central Europe.

Once widespread in Europe, Slender-billed Curlews were last observed in Brittany, France, in February 1968, and in North Yemen in January 1984. A colony, thought to be the last, was known on the Atlantic coast of Morocco in the 1990s, with the last reported sighting in the winter of 1997/8, although a flock of Slender-billed Curlews was photographed in southern Italy in  March 1995. No subsequent observations of the species have been made, despite extensive searches across its former range, including the Middle East and Central Asia.

The last known photograph of a group of Slender-billed Curlews, taken in southern Italy in March 1995. Marco Basso in van den Berg (1995).

The Slender-billed Curlew was first observed breeding in 1912 by Russian ornithologist Valentin Ushakov, who first noted that the species appeared to be in decline. The possibility that Slender-billed Curlews might be at risk of extinction was first raised in 1943 by German ornithologists Erwin Stresemann and Hermann Grote, but it was not until 1988 that the species was identified as Threatened by the International Union for the Conservation of Nature. The species was listed as Critically Endangered in 1994, following a re-assessment of its status, and an action plan for its recovery was produced in 1996. 

Based upon this long absence from its known range, and the extensive, yet unproductive, searched which have been made in the last three decades, combined with the declining number of sightings recorded throughout the twentieth century, Buchanan et al. carried out a statistical analysis of the probability that the species might still exist. Based upon this, they conclude that is most likely that the species became extinct in the 1990s, with only a miniscule possibility that the species might still exist in the 2020s.

If this analysis is correct, then the Slender-billed Curlew is only the third species of Bird known to spend a large part of its annual cycle in the Western Palaearctic to have become extinct since 1500, joining the Great Auk, Pinguinus impennis, last reported in 1844, and the Canarian Oystercatcher, Haematopus meadewaldoi, last observed in 1913. 

A preserved museum specimen of a juvenile male Slender-billed Curlew, Numenius tenuirostris. Naturalis Biodiversity Center/Wikimedia Commons.

It is difficult to directly assess the cause of the extinction of the Slender-billed Curlew, since it is likely that the species finally died out around the time that an action plan for its survival was first drawn up. When that plan was produced, it identified that the species was threatened by habitat loss across its range, and potentially being hunted in some areas as well. 

The only records of breeding by the Slender-billed Curlew are those made by Valentin Ushakov in southern Russia in the early twentieth century. It is possibly that the main breeding area for the species was further south, on the steppes of northern Kazakhstan, which were extensively converted to croplands by the Russian Empire and Soviet Union in the nineteenth and twentieth centuries, in the process destroying extensive wetlands, which might have served as breeding grounds for Slender-billed Curlews, however, without further evidence, this is purely speculative. 

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Saturday, 16 March 2024

Understanding the giant handaxes of the British Lower Palaeolithic.

The Lower Palaeolithic is the earliest stage in the archaeological record from which stone tools are known, and while such ancient tools are rare, they are much more common than the remains of the Hominins which produced them, or any other remains of their activity, thus providing a key insight into the activities of our ancient ancestors. The first bifacial handaxes appear in the African archaeological record about 1.76 million years ago, and are considered to be a significant conceptual leap from the Oldowan core-and-flake technology which preceded them, although whether these axes had an entirely novel function or were a new technological approach to tasks which were already undertaken is unclear. One of the notable manifestations of this technology was the manufacture of 'giant' handaxes, notably larger than other examples of these tools, sufficiently large that archaeologists are divided as to whether they had any practical function at all.

It has been suggested that the purpose of these giant axes may have been social rather than practical, with the production of such tools representing a significant time investment, and therefore being an indicator of patience and commitment to a project. Giant, highly symmetrical handaxes from British sites such as Furze Platt and Shrub Hill have been used to support this hypothesis. An alternative suggestion is that the Hominins involved may have found the manufacture of such giant axes an aesthetically pleasing activity in its own righty, an early sign of artistic expression emerging from the purely functional.

In a paper published in the journal Antiquity on 4 March 2024, Luke Dale of the Department of Archaeology at Durham UniversityAaron Rawlinson, also of the Department of Archaeology at Durham University, and of the Department of Britain, Europe and Prehistory at the British MuseumPete Knowles and Frederick Foulds, again of the Department of Archaeology at Durham University, Nick Ashton, also of the Department of Britain, Europe and Prehistory at the British Museum, David Bridgland of the Department of Geography at Durham University, and Mark White, once again of the Department of Archaeology at Durham University, present the results of a study of giant handaxes from across the British Lower Palaeolithic.

Dale et al. note that symmetry is often cited as a key feature of giant handaxes, but that no previous study appears to have actually looked at whether these axes are actually more symmetrical than other axes from the same period, nor whether the style or frequency of these objects varies over time. 

All giant handaxes are associated with the Acheulean technology, which in Britain was present by Marine Isotope Stage 13, about 500 000 years ago, and may have appeared as early as Marine Isotope Stage 15, about 600 000 years ago, if recent dates for the Fordwich and the Bytham River terraces are correct. The global climate underwent a series of oscilations during the Middle Pleistocene (781 000-126 000 years ago), which in Britain manifested as a series of warm interglacials during which Hominins invaded and settled the landscape, interspersed with colder glacial intervals during which they disappeared. This pattern results in a series of distinct cultural intervals within the British Acheulean, providing a potential opportunity to detect discrete phases in the manufacture of giant handaxes in Britain.

Dale et al. obtained data on 4160 handaxes of all sizes, from 47 sited dating to between Marine Isotope Stage 15 and Marine Isotope Stage 9. For convenience, each (odd) Marine Isotope Stages was considered to represent a warm phase, as well as the cool phases bracketing it, thus Marine Isotope Stage 11 in fact refers to the Marine Isotope Stage 10-11-12 cool-warm-cool cycle; each cycle therefore represents about 100 000 years. The warm phases covered by Dale et al.'s study are MIS 15 (approximately 610 000 to 560 000 years ago), MIS 13 (approximately 524 000-474 000 years ago), MIS 11 (approximately 427 000 to 364 000 years ago), MIS 9 (approximately 328 000-301 000 years ago. Many of the sites in the study are well dated, but others represent chronologically mixed assemblages, such as Dunbridge in Hampshire, where the artefacts were recovered from dredged aggregates, have only been tentatively dated, such as Whitlingham in Norfolk, or are known to have been the subject of collecting biases, such as Furze Platt in Berkshire. Dale et al. recognise that these factors will have an impact on the reliability of their study, but reason that the size of their dataset should offset this. They also note that a further two giant handaxes were discovered at Frindsbury in Kent after the initial daft of their study was submitted, and that these are thought to date to MIS 9.

Handaxes were divided into six classes on the basis of their symmetry, with Class 1 representing near perfect symetry, and Class 6 indicating almost no symmetry.

Dale et al. measured the length of all the 4160 handaxes in their study, splitting them into four categories; small (less than 80 mm in length), average (80-150 mm in length), large (150-220 mm in length) and giant (more than 220 mm in length.

Forty two of the handaxes fell into the 'giant' category, of which 33 can be confidently assigned to the MIS 9 cycle, the final stage of the Acheulian in Britain. A single giant axe was dated to MIS 11, and two to MIS 13 - although one of these is- derived from a MIS 6 gravel bed, and is interpreted as having been reworked from a MIS 13 deposit. Since MIS 9 only represents about 1% of the British Pleistocene record, this concentration of giant handaxes in MIS 9 deposits is interpreted as being significant.

In the 1960s archaeologist Derek Roe developed a system of classifying handaxes upon their shape. The majority of giant handaxes in Dale et al.'s study conform to Roe's Group I (pointed, with cleavers) and Group III (plano-convex), both of which are associated with MIS 9. Notably, handaxes in an assemblage from Broom on the Devon/Dorset border, which included several giant handaxes, while still from MIS 9, did not easily fit into Roe's scheme. The stone tools here were made from chert derived from the Upper Greensand, and tended to be ovate or asymmetrical, unlike other MIS 9 assemblages.

Other than the presence of giant handaxes in the MIS 9 material, the size distribution of handaxes between different interglacials varied little, with the average size of the axes varying only by 10-20%, although the MIS 15 handaxes appear on average slightly larger than those from later assemblages, and the MIS 11 assemblage contained a higher proportion of small handaxes than other stages. However, Dale et al. note that their MIS 15 material came from only two sites, Brandon Fields in Suffolk and Fordwich in Kent, with the material used at Fordwich being elongated pipeflint, which lends itself to the making of larger bifaced tools with a minimal amount of effort. Given this, Dale et al. are reluctant to read much into this data without further MIS 15 material becoming available.

Analysis also suggests that axes from MIS 13 and MIS 9 are on average larger than those from MIS 11, and MIS 9 axes are also slightly larger, on average, than those from MIS 13. Dale et al. are uncertain whether this variation relates to cultural differences between the different populations colonising Britain in each interglacial, and make no further assessment of this data at this time.

A comparison of handaxe length between MIS 15, 13, 11 and 9, with handaxes grouped according to length class. MIS 15 displays a greater proportion of handaxes in the large class, while small handaxes are much more limited. MIS 11 displays a greater proportion of small handaxes. Giant handaxes represent a very small percentage of the total assemblage, emphasising their status as extreme outliers beyond the usual variation in handaxe length. Frederick Foulds & Mark White in Dale et al. (2024).

The majority of giant handaxes come from the Thames Valley or its southern tributaries, although examples are also known from Norfolk Yare, the Axe Valley and Solent River deposits. The majority of sites where they are found are close to primary chalk-flint outcrops, where large flint nodules were available. However, some were found in flint-poor areas, such as Wolvercote and Stanton Harcourt in Oxfordshire, whereas older archaeological sites, such as Boxgrove in West Sussex, and Elveden in Suffolk, are located near excellent sources of large flint nodules, but yield no giant handaxes. 

No giant handaxes available from Stoke Newington in London, one of the UK's best MIS 9 sites, although it is known that the site did yield such axes; as the original excavator of the site, Victorian archaeologist Worthington Smith, illustrated several giant handaxes in his book, Man, the primeval savage, but then gave these away to distinguished visitors to the site; these axes are thought to be either in private collections or lost. 

Geological map displaying the locations of sites with giant handaxes in relation to underlying bedrock geology, coloured as per the British Geological Society scheme. Colours for chalk bedrock are highlighted in the legend. Note that large clasts in overlying superficial deposits largely reflect local geology and ‘exotic’ raw materials would generally be smaller in size. Dale et al. (2024).

Of the forty two giant handaxes in the study, thirty five (or 83%) are pointed, typically conforming to the lanceolate (lance-shaped) or ficron (having a pointed planform shape with biconcave edges) type. These shapes are considered to be part of the same typological grouping as chisel-ended cleavers, a grouping which is thought by many archaeologists to be restricted to MIS 9. Although ficrons are associated with giant axes, not all ficron handaxes are giant; known ficrons range in size from 63 mm to 307 mm.

Examples of giant pointed and ficron handaxes: (A) the ‘Beast of Biddenham’ (length 253 mm); (B) a giant ficron from Canterbury West (length 285mm) (photographs courtesy of the British Museum (A) and The Seaside Museum, Herne Bay (B)). Dale et al. (2024).

Since the overwhelming majority of the giant axes date from MIS 9, Dale et al. examined axes from this stage for their symmetry. Handaxes from this interval were found to vary a great deal in their degree of symmetry, although there was a direct corelation between axe length and symetry, with longer axes tending to be more symmetrical than shorter ones. 

Splitting the axes into groups based upon their size emphasised this trend, with axes in larger categories being more symmetrical than those in smaller size categories. Examination of earlier axes suggested that this tendency for larger axes to be more symmetrical was also present in MIS 11, but not MIS 15 or 13, although the number of axes available from the earlier periods makes this result less reliable. 

However, this tendency for larger axes to be more symmetrical in MIS 11 and 9 deposits may be influenced by another trend, which is for axes in general to be much more symmetrical in MIS 13 and 11, compared to other periods, and in particular to MIS 9. Thus, the increased symmetry seen in the large MIS 9 axes is balanced by a decrease in the symmetry seen in smaller axes, with the smallest axes (less thn 100 mm in length) tending to be roughly and quickly made, with very low symmetry. 

Giant handaxes are often cited as a key feature of Hominin cognitive and cultural development. Dale et al.'s study demonstrates that these items became prevalent in Britain during MIS 9, i.e. immediately prior to the Lower-Middle Palaeolithic transition, 300 000 and 250 000 years ago. This interval has already been identified as one of significant technological innovation, beginning with a non-handaxe phase, which was then followed by a period with handaxes, then a final phase in which a hierachical core-working technology, known as the proto-Levallois was introduced. The linking of giant handaxes to this stage adds another element to this increasing cultural complexity.

The presence of giant handaxes is often independent of the availability of suitable manufacturing materials, with giant handaxes known from areas of Oxfordshire where there are no sources of suitable large flints. This implies that either the raw material or the finished axes were transported for at least 25-40 km to the sites where they were found. This phenomenon has also been observed in France, with a giant handaxe found at Caune de l’Arago in Pyrénées-Orientales Department, more than 30 km from any suitable source of material. This transportation of materials is another new innovation from this phase, with most previous Lower Palaeolithic sites using stone sourced less than 5 km from the location where it was discovered. The increased effort being put into moving giant axes to locations away from the materials from which they were made lends further support to the hypothesis that these were valuable items, although another possibility is that these tools had a long working life, and were subject to regular reworking to maintain a sharp edge, which would explain the plano-convex form of many handaxes, including those at Wolvercote in Oxfordshire. Under this hypothesis, many large handaxes would have been discarded prematurely, before being extensively reduced.

Many of the giant handaxes are ficrons, a type of axe otherwise rare in Britain. Large ficrons are particularly striking objects, apparently crafted with care to maintain the length of the axe while reducing the width of the upper and central portions of the axe. This may have been a way to correct for a lack of symmetry in the original platform (the rock from which the ficron was made), or a sign of the tool having an extended lifespan, with regular resharpening. Another hypothesis is that the width reduction may have been a way to reduce the weight of the tool part of a general Acheulian trend for longer tools to be narrower. The alternative is that the ficron-shape was desired from the outset, and that the extra effort put into producing an axe of this shape is an indicator of the significance of the object to the people who made it. Whatever the truth, there does appear to be a link between the high symetry, large size, and unusual shape of these giant ficrons which appears to show a degree of artistry transcending the merely functional.

It has often been speculated that giant handaxes had no practical function, but experimentation has shown that examples far larger than anything found in Britain can be used as tools. Even the largest giant axes can be used two handed, can be used for digging, or can be used as static cutting tools across which materials could be drawn. Despite this possible functionality, all the British giant handaxes are distinctively formed and highly symmetrical, making it hard to believe that aesthetic considerations played no part in their manufacture. 

The oldest known giant axe is 1.6 million years old, and came from the Olduvai FLK West site in Tanzania. This is long before anything else considered to be even possibly an expression of communal artistic standards, and therefore was presumably made by an individual for reasons that were important to them alone. A more communal approach to aesthetics begins to appear about 500 000 years ago, at which point tools also being to appear which have been shaped to make the most of features such as fossils on the rock cortex. Giant handaxes may also represent an aesthetic development, with the size and symmetry of stone tools being increased to create visually pleasing objects. This implies the people who made these tools were capable in taking pleasure in creating such objects, which would imply an increasing level of cognitive and social development.

Giant handaxes may also have had some symbolic purpose. The manufacture and then apparent discarding of objects which take a degree of forward planning and labour investment may have been a way of marking territory, or signposting important locations to other members of the community.

The ability to produce giant handaxes clearly predates MIS 9; they are known from earlier in the Lower Palaeolithic and across the Acheulean world. A 300 mm-long lanceolate giant handaxe from Caune de l’Arago is thought to date from MIS 14, and other examples from across northern France, at sites including St Acheul and Thennes in the Somme Valley, Vaudricourt in Pas-de-Calais, and Montguillain in Oise Department, have variously been dated to MIS 11 and 9, although none of these dates are considered to be particularly secure. Giant handaxes are also known from the Levant and Middle East, though these tend to be less accurately dated. A 220 mm long example from Qesem Cave in Israel is thought to be between 420 000 and 200 000 years old, while a 265 mm-long basalt handaxe was found at Wadi Dabsa in Saudi Arabia. A site at Porto Maior in north-west Spain was occupied for a long interval across MIS 8-7, by a people of apparent African affinities, who regularly produced giant handaxes, picks and cleavers. Giant handaxes have been found at 23 sites across Africa, including Olduvai in Tanzania, where the largest such axe is 330 mm long and 1.4 million years old, Isenya in Kenya, where the largest axe is 253 mm long and 700 000 years old, and Kalambo Falls in Zambia, where the largest axe is 350 mm long and 300 years old. However, the African Acheulian, with which many sites in southern Europe and the Middle East are also associated, has a greater emphasis on large cleavers, picks, and handaxes, together referred to as 'large cutting tools'.

The MIS 9 giant handaxes from Britain appear to show a different phenomenon, a discrete technological interval in which larger, more symmetrical tools suddenly appeared. In France, the MIS 9 sites in the Somme Valley are the closest available to compare, with the Cagny l’Epinette locality producing 33 handaxes, the largest of which is still less than 170 mm in length, while the Revelles site has produced 52 handaxes, none more that 190 mm long. Ferme de l’Epinette, a MIS 10 site in the Somme Valley has produced 31 handaxes, all less than 180 mm long, while the MIS 9 stage of the Soucy complex in the Paris Basin has produced 178 handaxes, the largest of which are 155 mm long, and no handaxe from the entire complex exceeds 180 mm. The Orgnac Cave site in the Ardèche Department has a sequence of archaeological layers running from MIS 10 to MIS 8, but has produced no handaxes more than 200 mm in length. The MIS 9 layers at Menez-Dregan in Brittany have produced 29 handaxes, all less than 160 mm in length, although this may be because the tools are made from sandstone and microgranite beach pebbles which would not lend themselves to making bigger tools. This lack of large rocks for use as a source of tools may also account for the generally small handaxes found at Atapuerca in Spain.

Thus the production of giant handaxes in Britain appears to be a geographically, as well as chronologically, restricted phenomenon. This fits in with a more modern understanding of the Acheulian Culture, which includes many such small-scale variations, rather than being a single long-lived and widespread monoculture, as was once perceived. 

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