Showing posts with label Cervidae. Show all posts
Showing posts with label Cervidae. Show all posts

Tuesday, 29 August 2023

Archaeologists uncover ritually buried Red Deer from Bronze Age England.

Archaeologists carrying out exploratory work ahead of the construction of a new water supply network in the east of England have uncovered what appears to be a pair of ritually buried Red Deer at Navenby in Lincolnshire, according to a press release issued by Anglian Water on 9 August 2023. The Deer, which are thought to have been buried about 4000 years ago, were accompanied by pottery associated with the Bell Beaker Culture, a Bronze Age European Culture with appeared about 4800 years ago in Iberia, and spread across Europe and parts of North Africa, reaching Britain about 4450 years ago. The Deer show no signs of butchery (the removal of meat for consumption, which leaves cut marks upon the bones), and were therefore presumably intact when buried, although it is possible that they were ritually sacrificed for some purpose.

Remnants of two Red Deer found in a Bronze Age burrial at Navenby in Lincolnshire, England. Anglian Water.

Other than the burred Deer, no further traces of Bronze Age activity were found at Navenby, although the archaeologists did uncover a small Iron Age settlement, comprising two roundhouses and five smaller structures, which may have been grain stores, as well as the burial of three cremated and one uncremated individuals, which may date from the Roman period.

See also...

Follow Sciency Thoughts on Facebook.

Follow Sciency Thoughts on Twitter.


Saturday, 7 January 2023

Understanding the message behind Europe's Upper Palaeolithic cave art.

Prior to about 37 000 years ago, cave art in Europe consisted largely of patterns of hand-prints, rectangles, and dots. After this time, a wealth of engraved and painted images swept across the continent, depicting the Animals of the Late Pleistocene with a realism that still impresses modern artists. Almost all of these artworks depict large herbivorous Animals, thought to have been important food sources for hunters on the Eurasian Steppes. In most cases it is easily possible to tell what species is being depicted, and even what time of year the image was portraying. The paintings on the walls of the Lascaux Cave in southwest France show a sequence of rutting Animals which can be interpreted as an ethological calendar (calendar of Animal behaviour), and at other sites the presence of Cervids (Deer) with antlers, and the aggressive postures of these Animals, have been interpreted as depiction of a particular season.

However, this cave art does not just depict Animals, alongside these images are frequent abstract marks, mostly dots and lines, but including other shapes such as 'Y's. These marks are usually either directly adjacent to the Animal images, or actually overlie them, suggesting a strong connection between the two. Similar marks have been found carved onto bones, dating from the onset of the European Upper Palaeolithic (roughly equivalent to the African Middle Stone Age) onwards, and are generally thought to have had a recording function. It has been generally assumed that such marks are numeric, and probably recording periods of time, but their exact message has remained unclear. 

Previous studies have found that 66% of the Animal depictions in Upper Palaeolithic European cave art are associated with other symbols, although these vary in form, and probably convey a variety of messages. It is also likely, given the long chronological range and wide geographical area over which these symbols are found, that more than one form of symbolic recording is present. 

In a paper published in the Cambridge Archaeological Journal on 5 January 2023, independent researchers Bennett Bacon, Azadeh Khatiri, and James Palmer, together with Tony Freeth of the Department of Mechanical Engineering at University College LondonPaul Pettitt of the Department of Archaeology at the University of Durham, and Robert Kentridge of the Department of Psychology at the University of Durham and the Canadian Institute for Advanced Research programme in Brain, Mind and Consciousness, focus on two clear and simple patterns found in many of these depictions: Animals found associated with rows of dots and/or lines (which are presumed to be functionally the same) and branching 'Y' symbols. 

These symbols are found throughout the European Upper Palaeolithic, although most common from the later portion of this period, possibly implying that they became more common over time. Bennet et al. consider the meaning of the Animal depictions to be unambiguous, but consider the lines, dots, and 'Y' symbols to be potentially interpretable, and related to the ethology of the Animals they are found close to. The basis for this assumption is that the dots and lines in these sequences represent numbers, and that the combination of numbers and recognisable Animals gives the basis for deciphering messages tens of thousands of years ago.

Examples of animal depictions associated with sequences of dots/lines. (a) Aurochs: Lascaux, late period; (b) Aurochs: La Pasiega, late; (c) Horse: Chauvet, late (we differ in opinion with the Chauvet team, for whom it would be early); (d) Horse: Mayenne-Sciences, early; (e) Red Deer: Lascaux, late; (f) Salmon: Abri du Poisson, early; (g) Salmon (?): Pindal, late; (h) Mammoth: Pindal, early. Bacon et al. (2022).

It is now generally accepted that the regular marks left by Upper Palaeolithic peoples represent a form of counting, and therefore the storage of information outside of the brain. The processing of numbers can be done in several ways; by subitisation, or simply recognising small groups of objects at a glance (e.g. six tins in a box), by cardinality, or counting objects (twenty sheep in a field), or by ordinality, being able to place objects within a sequence (the third house in a row). Experimentally these have been shown to be distinct skills, carried out in different parts of the brain, and children have been shown to develop these skills in order as they develop, starting with subitisation and progressing to ordinality. 

In 1991, Alexander Marshack suggested that the dots and lines seen in Upper Palaeolithic art were numbers, most likely representing periods of time. In his view, each dot or line was likely to represent a day, with the sequences of pictures they were associated with being lunar calendars. The lunar calendar theory failed to win many supporters within the academic community, but the concept of the representations as a simple way of recording events outside the mind did gain wider acceptance. 

The grouping of engraved marks on bones and antlers from the Upper Palaeolithic has led to the conclusion that these marks were being used to store information, and that this information was most likely numeric. This has in turn led to the suggestion by cognitive archaeologist Karenleigh Overmann  that once Humans had begun to collect numerical data, it was almost inevitable that they would apply this to time, and in particular the seasons, important phenomena in the lives of people so closely dependent on nature.

The sequences of dots and lines found adjacent to Palaeolithic Animal images fit the same criteria for representing numbers as the carvings on bones and antlers, and are consistently arranged in lines horizontal to the orientation of the Animal images (which are often on ceilings). These sequences vary in length, but appear to have been recording numbers using the additive principle, where an extra mark represents an increase by one (as in the Roman numerals, I, II, III; or the Chinese 一, 二, 三).

In the 1990s archaeologist Carole Fritz assembled a database of 90 symbols used on portable objects associated with the Late Palaeolithic Magdalenian Culture, collected from the Dordogne and Pyranees, and including linear marks, dashes, angular signs, arc shapes, broken lines, dots, various impact marks and combinations and repetitions thereof. These symbols were used consistently at sites associated with the Magdalenian Culture, with no discernible variations over space and time. Based upon this, Fritz concluded that these marks represented a method of storing information recognised across the Magdalenian Culture, although the exact meaning implied remained obscure.

Hunting appears to have been an important source of nutrition to Upper Palaeolithic peoples, with the most important prey species being Horses, Bovids, Cervids, Caprids, Proboscideans, Birds, and aquatic Animals. Modern populations of these Animals show strong seasonal patterns of behaviour, with cycles of mating and birthing, and seasonal migrations in the spring and autumn. During this seasonal cycle, many of these Animals shift between large and small groups, and are present on different parts of the landscape.

Zooarchaeological studies have revealed a similar seasonality in the behaviour of these Animals during the Pleistocene, and it seems likely that a knowledge of migration patterns and breeding cycles would have been key information to Upper Palaeolithic hunters dependent on these Animals for survival. As such, it is unsurprising that these Animals were a central preoccupation for these people, and that images of them dominate their art. Furthermore, at sites such as Lascaux Cave depict these Animals in such detail that their pelage, hair, antler growth, gregarious and aggressive behaviour and other indications of rutting can be used to show that their breeding behaviour was also of great interest. Many of the Animal images at Lascaux are associated with symbols; for example a row of swimming Stags, generally interpreted as an autumn migration scene, is accompanied by a row of seven red dots, while a mating scene with male and female Aurochs in their summer coat is marked with four black dots. At a scene from the slightly later Font de Gaume Cave two Reindeer stags are depicted with locked antlers, and marked with a set of eight dots.

Bacon et al. accept the premise that the sequences of dots and lines associated with the Upper Palaeolithic Animal images are saying something about the Animals in the images with which they are associated. Furthermore, they accept that these are abstract representations, rather than parts of the images, noting that several different taxa are consistently annotated with the same number of dots, but on different parts of their anatomies. Given this consistency, they also accept that the message conveyed by the dots is numerical in nature, being either cardinal or ordinal.

Working from this, they assume that the objective was to convey some form of useful numeric information about specific prey Animals to future readers. Given this, it seems unlikely that they were recording the number of Animals sighted, the time at which they were sighted, or even the number of Animals killed. It is likely, however, that they would have wanted to record information about migrations, times when Animals could be relied upon to form aggregations, and times when Animals were most vulnerable to attacks by hunters (i.e. mating and birthing). Working from these assumptions, Bacon et al. propose that the timing of such events is the most logical numerical information to have been juxtaposed with the Animal images.

If Upper Palaeolithic people were able to record numbers using the additive principle, then it is quite likely that the rows of dots or lines seen on their art represent amounts of time. Furthermore, the number of dots or lines associated with an Animal image is never particularly high (and never higher than thirteen), so if each dot or line represents a single unit of time, then that unit is almost certainly a lunar month, since this is one of the few units which would have been obvious to pre-agricultural people. Based upon this, Bacon et al. believe that the dots and lines present in Upper Palaeolithic images represent lunar months, presenting information about the behaviour of Animals on a seasonal calendar. 

If this is correct, then the calendar being used by these people must have had an established beginning; a time of year against which the tallies could be counted, giving the people using it a point to count from. Most historical calendars have their roots in astronomical observations, counting from the solstices and equinoxes. These give accurate dating systems usable by agriculturalists, but are difficult to observe, and likely to have been of limited use to Palaeolithic hunters. Rather, these people lived in a world dominated by meteorological events, and seasonal cycles in temperature and the behaviour of Plants and Animals. One seasonal event which Bacon et al. believe would have been hard for Palaeolithic peoples to miss would have been the annual spring thaw. This is widely referred to using the French term bonne saison by zooarchaeologists, marking a period when rivers unfroze, snow melted, and the world began to turn green. This would not have occurred everywhere at the same time, with spring coming several weeks earlier in southern Europe than in the north, but would have provided a useful starting point for a yearly calendar used by people needing to track the migrations and mating habits of Animals, but not exchange accurate date information with distant communities. 

One of the problems for peoples wanting to record time in the ancient world was that the solar year cannot be divided into an exact number of lunar months (there are approximately 12.37 lunar months in one solar year). This led to the development of a variety of complex calendars, with periodic adjustments being made to bring the solar and lunar cycles into alignment. None of this is likely to have mattered much to Upper Palaeolithic peoples counting from the bonne saison, who could have just counted the lunar months from each spring thaw into the depths of winter, when it would have become irrelevant. 

Bacon et al. therefore theorise that the sequences of dots and lines in Upper Palaeolithic art represent lunar months after the bonne saison. Therefore, if an Animal is represented along with a sequence of three dots, then the intent was to imply some important feature of that Animal's life cycle happened three months after the spring thaw. This is technically an interval calendar, recording that events happened a certain period after the bon saison, rather than a true calendar recording accurate dates.

Furthermore, Bacon et al. note that many of the sequences contain a 'Y' symbol in the midst of a series of dots or lines, which they believe represents a significant event in the life cycle of these Animals, and reason must be one of four possible annual occurrences, spring migration, mating, birthing, or autumn migration. 

The 'Y' symbol is one of the most commonly depicted symbols in Upper Palaeolithic art, and therefore presumably has an important meaning. Bacon et al. note that the position of this symbol within sequences varies between Animal taxa, but remains constant for each taxon, implying that it is carrying species specific data, and that that data appears to be ordinal in nature, within an ordinal sequence. Thus the total number of symbols potentially records one piece of information, with the position of the 'Y' symbol conveying a second. Bacon et al. further propose that, given the known biology of the Animals involved, the 'Y' symbol is most likely to represent birthing.

In order to test this hypothesis Bacon et al. compiled a database of Animal symbols with associated symbol sequences, and compared these to known data on the reproduction and migration of their modern relatives, and information about Pleistocene Animals established by previous zooarchaeological studies. Having eliminated any dubious or ambivalent sequences from the record, Bacon et al. were left with 256 sequences containing a 'Y' symbol and 606 sequences which did not.  The majority of these sequences came from France and Spain, although some came from further east. Chronologically, the images span the whole of the Late Upper Palaeolithic, although the majority come from the end of this period. 

Bacon et al. initially divided their data into two chronological sections, the Early and Middle Upper Palaeolithic Aurignacian and Gravettian cultures, and the Late Upper Palaeolithic Solutrian and Magdalenian cultures, but surprisingly no difference could be found between these datasets, suggesting that, contrary to expectations, the same data-recording system seems to have persisted in Europe for over 20 000 years. 

For the sake of convenience, Bacon et al. sorted some of the Animals depicted into groups, such as Cervids, Caprids, Birds, and Fish, while others were retained as separate species, i.e. Bison, Aurochs, Horses, Mammoths and Rhinos. Other species, such as Snakes and Wolverines, were present, but in very low numbers, and were excluded from the dataset, as were images of Humans.

In order to make the comparison, Bacon et al. converted the expected birthing season for the Animals into months relative to the Pleistocene bonne saison. Thus, assuming the bonne saison started around the start of May, the birthing season for European Bison, typically August, should be 3 or 4. Such a calendar is necessarily approximate, given that it is based upon seasonal events which might themselves vary from year to year, and will certainly vary regionally.

Bacon et al.'s prediction was that the 'Y' symbols related to each Animal group should be clustered, rather than randomly distributed throughout the sequences, and that these clusters should correspond to the predicted birthing seasons of the Animals, adjusted to an ordinal calendar rooted at the bonne saison. Since this should be different for the different groups, the position of the 'Y' symbol in the sequence should also vary between groups.

This prediction proved to be true for the majority of the groups, with the 'Y' symbol being both consistent and coinciding with the predicted birthing season for Aurochs, Bison, Horses, Fand Mammoths, and less clear but still matching the prediction for Cervids. Only Caprids failed to match the prediction, with no clear pattern observable in the data for this group. No correlation could be made between the position of the 'Y' symbols and the predicted times of mating or migration. However, the observed total number of marks correlated to the predicted mating season for Aurochs, Bison, Horses, Mammoths, and probably Cervids.

The two non-Mammalian groups, Birds and Fish, showed slightly different patterns, with the 'Y' correlating with mating and the end of the line with hatching for Birds, and the 'Y' correlating with the spring migration and the end of the line with hatching in Fish; in both cases the sequence relates to the appearance of a food source.

Bacon et al.'s data provides strong statistical evidence for a correlation between the position of a 'Y' symbol in a sequence associated with an image of an Animal, and the birthing season of that Animal, as calculated on an interval lunar calendar rooted in the Pleistocene bonne saison. It also provides weaker statistical support for the total number of marks representing the mating season of the same Animals. 

The idea of Upper Palaeolithic peoples in Europe using a form of numeric notation using notches, lines, and other marks has been accepted for some time, and is no longer really controversial. Based upon this, it has also been generally assumed that other marks used by these people were recording some form of information, although the exact nature of this information has, until now, remained unclear. 

Bacon et al. propose that the symbols associated with the (unambivalent) Animal images relates directly to the biology and behaviour of those Animals, providing a key to understanding the Upper Palaeolithic notation system. This is based on the assumption that the position of a particular mark, the 'Y' symbol, within a sequence of simpler marks (dots or lines), represents a key event in the life cycle of these organisms, in this instance the birthing season, something supported by a statistical analysis of the data.

This means that these Upper Palaeolithic people had a means of preserving information which could be read back after thousands of years, by people who might have quite different languages, but understood the common notation system (we know little of the languages spoken in the Upper Palaeolithic, but they are unlikely to have remained constant for tens of thousands of years). The widespread use of this system over long periods of time and a wide geographical area suggests that it was important to record this information in a way that went beyond oral traditions (although these are likely to have also been used).

The symbols represent more than a simple tally, and Bacon et al. propose that they represent the development of a simple calendar, based upon meteorological events and the biology of important prey Animals. This would predate by many thousands of years any previously known calendar or writing system.

It is far from clear how widely the interpretation of this data would have been understood in the communities which produced it. Objects, such as bones or antlers, with tallies on them are widely known from this period, and would presumably have been present in domestic environments, carried by members of the community. However, the cave art was typically placed in deep cave environments, where it was potentially not seen by the whole community, and may have been restricted to a small number of people.

Bacon et al. do not argue that their interpretation is exclusive of the artworks having had other purposes, aesthetic or ritual, nor do they claim to have unravelled all of the meaning behind the symbols (other symbols are present in the cave art, which they have not included in this study). Furthermore, they do not assert that this notation system was used by all peoples present in Europe throughout the Upper Palaeolithic, and acknowledge that there are plenty of instances of Upper Palaeolithic art in which Animals are depicted without the presence of additional symbols of any kind.

They do however, believe they have uncovered a method of storing data about the behaviour of important prey Animals, which was used in parts of western and central Europe for a period which lasted from about 37 000 years ago to about 13 000 years ago. In this interpretation, the individual images do not represent individual Animals, but are instead representative of entire species, and the behaviour of these species as experienced by the creators of the art. 

The rows of symbols and position of marks within them represent a simple syntax combining linear and ordinal numbers to record and communicate information, a form of intellectual abstraction, something which is considered to a key achievement of modern Humans (Upper Palaeolithic peoples are universally accepted as fully Modern Humans). This ability to record a combination of information derived the behaviour of Animals, number-based information, seasonal meteorology, and lunar months, therefore represents a significant intellectual achievement. This provided a system to preserve this information that went beyond oral tradition, and enabled the comparison of data from multiple years, presumably enabling the people who collected it to make estimations about variations in natural phenomena, something hard to do with purely oral records.

This raises the question of whether this system could genuinely be called writing. The system does seem to be able to deal with discrete quantities, i.e. using numbers to say something about the Animals they are associated with, rather than just counting the numbers. Furthermore, the use of placement to determine the value of the 'Y' symbol is a precursor of the use of place to show the value of a number (as in 1, 10, 100), something previously assumed to have been a Sumerian invention. It could even be considered that the 'Y' symbol represents a verb, 'to give birth', although this is less than clear, it could simply be a noun such as  'birth' or 'birthplace'.

The common modern use of the term language implies that it has a phonetic connection to a spoken language, generally that used by the writer. This is again thought to have originated in ancient Sumer, Mesopotamia, around 3300 BC, when a system comprising a mixture of pictograms and abstract numerical symbols first appeared. These system evolved into Cuneiform, which is considered to be a script rather than a language (in the sense that the modern English and French languages are written in Latin script). A form of record-keeping using small tokens is known to have been used in the Near East in the Neolithic, during the tenth millennium BC, with the system evolving over time and being widespread in the region by the sixth millennium BC. By the fourth millennium BC these tokens appeared to have taken other, non-numeric functions, something which evolved further into the Cuneiform script of Uruk-era Sumer. 

Bacon et al. postulate that the European Upper Palaeolithic system actually conveys more information than the first Mesopotamian scripts, in that in relates the behaviour of wild Animals to a meteorologically derived calendar, whereas the earliest Mesopotamian records appear to have just recorded quantities of items. Based upon this, they argue that this system can justifiably be called a script. However, they note that the system gives no indication as to how the people using it would have described the Animals depicted, the Moon or its phases, or the bonne saison, although they do assume that these people would have been able to describe all these things orally.

The European Upper Palaeolithic recording system appears to contain no grammatical syntax, which would justify its description as a true written language, but does seem to be recording data at least as well as the proto-Cuneiform script of ancient Sumer (although, unlike the Sumerian records, this system could not be described as 'administrative documents'). 

For this reason Bacon et al. do not press the claim that this system should be seen as a language (something they see as potentially controversial, and at best semantic), but a form of proto-writing forming a step between a simple tallying system and a true writing system. They are aware that this claim is likely to be contested, and that others in the field are likely to come up with other ideas about what this system should be called. 

For the time being they restrict themselves to describing the system as a form of proto-writing, conveying a phrenological/meteorological calendar, although they welcome debate on the subject. However this system should be described, it clearly represents a system of symbolic record-keeping dating from tens of thousands of years before the earliest Sumerian writing system, making it a highly significant discovery.

See also...

Follow Sciency Thoughts on Facebook.

Follow Sciency Thoughts on Twitter.


Saturday, 21 December 2019

Opossum seen grooming Deer for Ticks in Vermont.

The Vermont Wildlife Coalition has released a camera-trap photograph in which as Virginia Opossum, Didelphis virginiana, can be seen removing Ticks from the face of a Deer, which has apparently approached the Marsupial for this service. Virginia Opossums are known to be major consumers of Ticks, and therefore thought to be a significant control of Tick-borne diseases such as Lyme disease, which they usually acquire by visiting Tick-infested areas of woodland and then picking off any Ticks that attempt to attach to them, but this is thought to be the first time another animal has been seen approaching an Opossum for grooming, a form of behaviour more associated with marine organisms such as Cleaner Fish.

A Virginia Opossum, Didelphis virginiana, grooming a Deer in a forest in Vermont. Vermont Wildlife Coalition.

Virginia Opossums are North America's only native Marsupial, and, unlike many Marsupials, are extremely adaptable in their habits, rapidly taking to new environments and food sources when these become available. They were confined to the southwestern United States, Mexico, and Central America as far as Costa Rica, until the early twentieth century, since when they have colonised much of the rest of the United Stares and southern Canada, spreading through man-made habitats, such as farms and gardens, into areas such as the forests of New England where they were not previously found.

See also...

https://sciencythoughts.blogspot.com/2019/10/hundreds-of-koalas-feared-to-have-died.htmlhttps://sciencythoughts.blogspot.com/2018/02/wakaleo-schouteni-new-species-of.html
https://sciencythoughts.blogspot.com/2016/10/gumardee-richi-gumardee-springae-two.htmlhttps://sciencythoughts.blogspot.com/2015/06/unexpected-social-behaviour-in-south.html
https://sciencythoughts.blogspot.com/2014/04/reconstructing-diet-of-miocene.htmlhttps://sciencythoughts.blogspot.com/2013/09/hunting-lost-opossum.html
Follow Sciency Thoughts on Facebook.

Thursday, 13 June 2019

Muntiacus gigas: A new specimen of the Giant Muntjac from the Early Holocene of northern Vietnam.

Muntjacs, Muntiacus spp., are small solitary Deer found across South, Southeast, and East Asia, though with their earliest fossils found in the Miocene of Central Europe. The genus currently contains about a dozen species, with several having only been described in the last few decades. The living Giant Muntjac was first described from northern Vietnam as Megamuntiacus vuquangensis, in 1994, being placed in a separate genus on account of its larger size, though this was later considered to be erroneous, as the species is not sufficiently genetically distinct to justify this. It was then realised that the species was identical to one described from Hemudu, a Middle Holocene (6000-7000 years old) Neolithic site in Zhejiang Province in eastern China, as Muntiacus gigas, in 1990, and then thought to represent an extinct Muntjac species. Since the Chinese specimen was named first, this name takes precedence, and is considered the valid name for the species. The species is currently confined to the Annamite Mountains of Vietnam, Laos, and Cambodia, with this range inferred mostly from hunting trophies in museum collections, and is currently considered to be Critically Endangered under the terms of the International Union for the Conservation of Nature's Red List of Threatened Species, with a high likelyhood of going extinct within the next 20 years. However, the presence of the species in the Middle Holocene of eastern China suggests that the species once had a much greater range.

In a paper published in the journal Royal Society Open Science on 13 March 2019, Christopher Stimpson of the School of Natural and Built Environment at Queen’s University Belfast, Benjamin Utting of the Department of Archaeology at the University of Cambridge, Shawn O’Donnell, also of the School of Natural and Built Environment at Queen’s University Belfast, Nguyen Huong of the Institute of Archaeology at the Vietnam Academy of Social Sciences, Thorsten Kahlert, again of the School of Natural and Built Environment at Queen’s University Belfast, Bui Manh of the Department of Tourism of Ninh Bình Province, Vietnam, Pham Khanh of the Tràng An Landscape Complex Management Board, and Ryan Rabett, once again of the School of Natural and Built Environment at Queen’s University Belfast, describe a new specimen of Muntiacus gigas from an Early Holocene archaeological site in the Tràng An World Heritage Area in Ninh Binh Province, northern Vietnam.

The Tràng An World Heritage Area lies on the southern margin of the Red River Delta in Ninh Binh Province, northern Vietnam, and comprises a karstified (eroded) limestone massif covered by forest that rises from the coastal plain. The Hang Boi Cave Complex is an archaeological site within this area, comprising a series of interconnecting caverns with a southeast-facing entrance, which shows signs of having been inhabited during the Late Pleistocene and Early Holocene,

The specimen in the study (HBC-27587) comes from a midden pile at the entrance to the cave, comprising largely Mollusc shells, but also bones from other Mammals, Birds and Turtles. The specimen is a partial mandible (jaw bone) identified on the basis of bone and tooth anatomy as belonging to Muntiacus gigas. The specimen has been dated to between 11 100 and 11 400 years old.

Specimen HBC-27587 shown in lateral (b), medial (c), dorsal (d ) views and occlusal surfaces of m2 and m3 (e). All scale bars ¼ 20 mm. Stimpson et al. (2019).

The presence of Muntiacus gigas in the Early Holocene of northern Vietnam is not surprising, given that the species is known to have ranged as far as eastern China in the past. However the environment at Hang Boi is very different to that occupied by the species today, suggesting that it has altered its habits in response to Human pressure; modern Giant Muntjacs are found in dense forests in the Annamite Mountains, sometimes at 1200 m above sealevel. However the Tràng An specimen, while it may have been carried a short distance by Humans before being deposited, appears to have lived in a very different environment, on a lowland plane at most 200 m above sealevel, which is interpreted as having been covered by an open Oak woodland during the early Holocene.

See also...

https://sciencythoughts.blogspot.com/2018/08/assessing-impact-of-large-animal.htmlhttps://sciencythoughts.blogspot.com/2016/08/lightning-kills-323-reindeer-in.html
https://sciencythoughts.blogspot.com/2016/01/animal-remains-from-middle-neolithic.htmlhttps://sciencythoughts.blogspot.com/2016/01/mapping-distribution-of-southern-pudu.html
https://sciencythoughts.blogspot.com/2015/12/capreolus-constantini-roe-deer-from.html
Follow Sciency Thoughts on Facebook.

Monday, 27 August 2018

Assessing the impact of large animal carcasses on plant biodiversity, using data from a Norwegian Reindeer herd killed by lightning.

A total of 323 Reindeer, Rangifer tarandus, were killed by a lightning strike during a thunder storm on the Hardanangervidda National Park on Friday 26 August 2016. The Reindeer migrate across the park, which is a mountainous plateau, at this time of year, and are believed to have huddled together during the storm, an action which helps to protect the younger members of the herd (which included 70 calves) from the worst of the elements, but which makes them particularly vulnerable to (rare) lightning strikes. While tragic for the Reindeer, this event provided a unique opportunity to study the impact of large cadavers on a (nutrient poor) tundra environment.

Reindeer killed by lightning on the Hardangervidda Plateau on 26 August 2016. Håvard Kjøntvedt/Norwegian Environment Agency.

In a paper published in the journal Biology Letters on 15 August 2018, Sam Steyaert of the Department of Natural Sciences and Environmental Health at the University of South-Eastern Norway, and the Faculty of Environmental Sciences and Natural Resource Management at the Norwegian University of Life Sciences, Shane Frank, also of the Department of Natural Sciences and Environmental Health at the University of South-Eastern Norway, Stefano Puliti, also of the Faculty of Environmental Sciences and Natural Resource Management at the Norwegian University of Life Sciences, and of the Norwegian Institute of Bioeconomy Research, Rudy Badia, again of the Faculty of Environmental Sciences and Natural Resource Management at the Norwegian University of Life Sciences, Mie Arnberg, again of the Department of Natural Sciences and Environmental Health at the University of South-Eastern Norway, Jack Beardsley of the Surveying and Spatial Sciences Group at the University of Tasmania, Asle Økelsrud, once again of the Department of Natural Sciences and Environmental Health at the University of South-Eastern Norway, and Rakel Blaalid of the Norwegian Institute for Nature Research, describe the results of a study which monitored the site of the 2016 mass Reindeer site for seeds introduced by scavengers, as a way of demonstrating that endozoochory (seed dispersal via ingestion by vertebrate animals) provides a viable means for plants to reach a new nutrient source within an otherwise nutrient poor environment.

The vegetation at the sire of the study was dominated by Dwarf Birch, Betula nana, as well as Heathers, Grasses, and an understory of Mosses and Lichens. A range of Mammals and Birds were seen scavenging at the site, including Corvids (Raven, Corvus corax, and Hooded Crow, Corvus cornix), Golden Eagle (Aquila chrysaetos), Foxes (Red Fox, Vulpes vulpes, and Arctic Fox, Vulpes lagopus), Wolverine (Gulo gulo) and several Rodents (Arvicolidae). In addition a number of small Birds were observed feeding on Blowflies (Calliphoridae) attracted to the carrion.

Steyaert et al. searched the site for feces of mesopredators, Birds and Rodents, finding a particular corelation between Bird feces and carcass density (i.e. there was more Bird erces in areas where the carcasses were most dense). They then examined the Bird feces for seeds of the Crowberry, Empetrum nigrum, keystone species of the alpine tundra with predominantly vegetative reproduction, finding 87.5% of the feces sampled contained viable seeds.

The Crowberry, Empetrum nigrum, an Arctic keystone species shown to be distributed to sites with carrion by scavenging Birds. Wikimedia Commons.

See also...

https://sciencythoughts.blogspot.com/2018/02/vultures-and-lions-poisoned-outside.htmlhttps://sciencythoughts.blogspot.com/2017/12/understanding-dispersal-of-rockwood-and.html
https://sciencythoughts.blogspot.com/2017/02/isaberrysaura-mollensis-neornithischian.htmlhttps://sciencythoughts.blogspot.com/2016/08/lightning-kills-323-reindeer-in.html
https://sciencythoughts.blogspot.com/2016/08/ceratocaryum-argenteum-plant-producing.htmlhttps://sciencythoughts.blogspot.com/2016/04/understanding-role-of-bears-in-enabling.html
Follow Sciency Thoughts on Facebook.

Saturday, 9 January 2016

Mapping the distribution of the Southern Pudú in Chile.

The Southern Pudú, Pudu pudu,  is one pf the world's smallest Deer species, typically weighing 35 to 45 cm at the shoulder and weighing 6.4 to 13.4 kg. It is found in southern Chile and Argentina, and is listed as Vulnerable on the International Union for the Conservation of Nature's Red List of Threatened Species, with an estimated population of about 10 000 individuals, and is known to be at threat due to habitat loss, competition with introduced species and predation by feral Dogs. However it is both small and secretive in nature, and its true distribution and numbers are therefore somewhat unclear.

In a paper published in the journal BMC Ecology on 7 January 2015, Melissa Pavez‑Fox of the Instituto de Ciencias Ambientales y Evolutivas at the Universidad Austral de Chile and the Programa de Magíster en Ciencias Biológicas mención Neurociencia at the Universidad de Valparaíso and Sergio Estay of the Instituto de Ciencias Ambientales y Evolutivas at the Universidad Austral de Chile and the Center of Applied Ecology and Sustainability at the Pontificia Universidad Católica de Chile describe a project to map the distribution of Southern Pudú in Chile using recorded observations of the species to establish its ecological tolerances.

A female Southern Pudú in the Los Lagos Region of Chile. Rodrigo Fernández/Wikimedia Commons.

Pavez-Fox and Estay collected data from 73 recorded occurrences of the Southern Pudú in Chile and 62 from Argentina to establish the ecological tolerances of the species. At each of these locations the average annual temperature, mean diurnal temperature range, temperature seasonality, maximum temperature of the warmest month, minimum temperature of the coldest month, annual precipitation, seasonal precipitation, precipitation over the wettest quarter, precipitation of the driest quarter, precipitation of the warmest quarter, precipitation of the coldest quarter and altitude was found and incorporated into the study.

Using this data a map of suitable territory for the Southern Pudú in Chile in Chile was established. It was found that most of the suitable territory was located at low altitudes within the Central Valley between Andes and the Cordillera de la Costa. Only 5.87% of this range was within protected areas, which were concentrated at higher altitudes.

(a) Projection of the model fitted for the Chilean territory. Colors represent the suitability of each pixel for Puda puda habitat. (b) Binary map of the projection of the model fitted for the Chilean territory (red) with respect to the distribution determined by the IUCN (dark grey) for Puda puda. (c) Overlap areas between Puda puda suitable areas and protected areas according to the model (red). Pavez-Fox & Estay (2015).

Pavez-Fox and Estay note that the distribution of the Southern Pudú is poorly covered by the Chilean National Parks system, which has been criticized on other occasions for being concentrated to much in the north of the country, despite much of the nation's distinctive biodiversity being found in the south. They further suggest that as a charismatic species the Pudú could be used as a 'flagship species' to help develop better protection for the distinctive habitats of southern Chile.

See also...

http://sciencythoughts.blogspot.co.uk/2016/01/pangolin-trading-in-mong-la-myanmar.htmlPangolin trading in Mong La, Myanmar. Pangolins, Pholidota, are highly specialized Mammals distantly related to the Carnivora (Dogs, Cats, Bears etc.). They are specialized Ant-eaters, lacking teeth and having long, sticky tongues used to extract the Insects from their nests. Uniquely among Mammals Pangolins are covered by keratin scales similar to those of...
http://sciencythoughts.blogspot.co.uk/2015/12/capreolus-constantini-roe-deer-from.htmlCapreolus constantini: A Roe Deer from the Early Pliocene of Hidalgo State, Mexico.   Deer, Cervidae, are one of the most abundant and diverse groups of Artiodactyls (even-toed...
http://sciencythoughts.blogspot.co.uk/2015/04/ancient-collagen-provides-insights-into.htmlAncient collagen provides insights into the relationships of the South American ‘Ungulates’.                                                                  South America was an isolated continent for much of the Cainozoic,  connected only to Antarctica during the earlier part of the period and...

Follow Sciency Thoughts on Facebook.


Sunday, 13 December 2015

Capreolus constantini: A Roe Deer from the Early Pliocene of Hidalgo State, Mexico.

Deer, Cervidae, are one of the most abundant and diverse groups of Artiodactyls (even-toed hoofed Mammals) today. They first appear in the fossil record in Eurasia in the Early Miocene, with the oldest specimens found in North America dating from the Early Pliocene on Florida. Today the Americas have a distinct Deer fauna of their own, comprising the genera Hippocamelus, Mazama, Blastocerus, Ozotoceros, Pudu and Odocoileus, with the Arctic Reindeer Rangifer tarandus and Elk Alces alces found in both the Old and New Worlds.

In a paper published in the journal Acta Palaeontologica Polonica on 16 April 2014, Eduardo Jiménez-Hidalgo of the Laboratorio de Paleobiología at the Universidad del Mar and Victor Bravo-Cuevas of the Museo de Paleontología at the Universidad Autónoma del Estado de Hidalgo describe a Roe Deer, Capreolus constantini from the Early Pliocene Atotonilco El Grande Formation of Hidalgo State in Central Mexico.

The specimen comprises a fragment of mandible with fragments several teeth, plus both humeri, part of an ulna, a tibia, a metacarpal, part of a metatarsal, the left patella, the first right phalanx of a forelimb and an isolated tooth. It was found 60 cm above a volcanic ash layer dated to 4.57 million years ago by fission-track dating and 4.2 million years ago by argon/argon isotope dating, implying a slightly younger age.

Mandible and teeth of the Cervid Mammal Capreolus constantini, from the Pliocene of Hidalgo, Mexico. (A) Left mandible fragment in lateral (A1) and occlusal (A2) views; partial p3 and p4 in occlusal view (A3); p4 and m1 in occlusal view (A4). (B) p3 in occlusal view. Jiménez-Hidalgo & Bravo-Cuevas (2015).

To date the earliest, Roe Deer, Capreolus spp., known from the fossil record was a specimen of Capreolus constantini (the same species as the Hidalgo specimen) known from the Late Pliocene of Udunga in the Republic of Buryatia in eastern Siberia. Both of the living members of the genus, Capreolus capreolus and Capreolus pygargus are also native to Eurasia and the group have generally been assumed to be exclusively Eurasian in origin and distribution, while the New World Deer are thought likely to have originated from a single colonisation event and subsequent diversification. The Hidalgo specimen is the oldest known Roe Deer fossil and of approximately the same age as the oldest known American Deer fossils. It also makes the Roe Deer the first known group of non-Arctic Deer to be found in both the Old and New Worlds. This implies that Deer made the transition between northeast Asia and North America at least twice during the Late Miocene/Early Pliocene, with either a population of Roe Deer making the crossing separately of the ancestors of modern American Deer, or Roe Deer being part of the American Deer group that subsequently crossed back into Asia.

See also...

http://sciencythoughts.blogspot.co.uk/2015/02/the-genetic-diversity-and-distribution.htmlThe genetic diversity and distribution of modern Giraffes.                                                        Giraffes, Giraffa camelopardalis, have traditionally been split into nine subspecies based upon their coat patterns, details of the skeleton and geographic distribution. Unlike many other large Mammals...
http://sciencythoughts.blogspot.co.uk/2014/07/a-new-species-of-early-eocene.htmlA new species of Early Eocene Artiodactyl named after Lady Gaga.                               The Artiodactyls (even-hooved Mammals) are the...
http://sciencythoughts.blogspot.co.uk/2014/07/a-highly-specialized-musk-ox-from-late.htmlA highly specialized Musk Ox from the Late Miocene of Gansu Province, China.              In 1932 Birger Bohlin, chief palaeontologist with the explorer Sven Hedin’s Sino-Swedish Scientific Expedition to Northwest China, discovered an...
Follow Sciency Thoughts on Facebook.